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Smithsonian National Museum of Natural History
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Department of Vertebrate Zoology

Division of Amphibians & Reptiles

Phyllomedusa vaillantii
Phyllomedusa vaillantii Ecuador, Pastaza Province. Photographed by William W. Lamar
bar Ronald Heyer
    W. Ronald Heyer
    Research Zoologist

  • Phone: (202) 633-0730
  • Fax: (202) 357-2986
  • E-mail: heyerr@si.edu

  • Mailing Address:
    Smithsonian Institution
    PO Box 37012, MRC 162
    Washington, DC 20013-7012

  • Shipping Address:
    Smithsonian Institution
    National Museum of Natural History
    10th & Constitution Ave NW
    Washington, DC 20560-0162

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Related Links

Complete List of Heyer's Publications and Downloadable PDFs
Frogs of Boracéia in PDF Format (Online Audio Supplement)
Leptodactylinae EndNote® Bibliography

Education

Ph.D.,University of Southern California, 1968, Biology.
M.S., University of Southern California, 1965, Biology.
B.A., Pacific Lutheran University, 1963, Biology.

Research Interests

Herpetology, Systematics, Evolution, Ecology; current focus is on systematics and biogeography of the Neotropical frog genus Leptodactylus.

Recent Publications

de Sá, Rafael O., Grant, Taran, Camargo, Arley, Heyer, W. R., Ponssa, Maria L. and Stanley, Edward 2014. Systematics of the Neotropical Genus Leptodactylus Fitzinger, 1826 (Anura: Leptodactylidae): Phylogeny, the Relevance of Non-molecular Evidence, and Species Accounts. South American Journal of Herpetology, 9: S1-S128. doi:10.2994/SAJH-D-13-00022.1

Heyer, W. R. 2014. Morphological analyses of frogs of the Leptodactylus latrans complex (Amphibia, Leptodactylidae) from selected localities in South America. Proceedings of the Biological Society of Washington, 126(4): 369-378. doi:10.2988/0006-324X-126.4.369

Heyer, W. R. 2014. Morphological analyses of frogs of the Leptodactylus latrans complex (Amphibia, Leptodactylidae) from selected localities in South America. Proceedings of the Biological Society of Washington, 126(4): 369-378. doi:10.2988/0006-324X-126.4.369

Heyer, W. R. and Heyer, Miriam M. 2013. Systematics, distribution, and bibliography of the frog Leptodactylus insularum Barbour, 1906 (Amphibia: Leptodactylidae). Proceedings of the Biological Society of Washington, 126(3): 204-233. doi:10.2988/0006-324X-126.3.204

Heyer, W. R. and Heyer, Miriam M. 2012. Systematics, distribution, and bibliography of the frog Leptodactylus validus Garman, 1888 (Amphibia: Leptodactylidae). Proceedings of the Biological Society of Washington, 125(3): 276-294. doi:10.2988/12-04.1

Heyer, Miriam M., Heyer, W. R. and de Sa, Rafael O. 2012. Leptodactylus pentadactylus. Catalogue of American Amphibians and Reptiles, 887: 1-48.

Heyer, W. R. and de Sá, Rafael O. 2011. Variation, Systematics, and Relationships of the Leptodactylus bolivianus Complex (Amphibia: Anura: Leptodactylidae). Smithsonian Contributions to Zoology, 635: 1-58. doi:10.5479/si.00810282.635.1

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