Related Links

Complete List of Heyer's Publications and Downloadable PDFs
Frogs of Boracéia in PDF Format (Online Audio Supplement)
Leptodactylinae EndNote® Bibliography

Education

Ph.D.,University of Southern California, 1968, Biology.
M.S., University of Southern California, 1965, Biology.
B.A., Pacific Lutheran University, 1963, Biology.

Research Interests

Herpetology, Systematics, Evolution, Ecology; current focus is on systematics and biogeography of the Neotropical frog genus Leptodactylus.

Recent Publications

Heyer, W. R. and Heyer, Miriam M. 2012. Systematics, distribution, and bibliography of the frog Leptodactylus validus Garman, 1888 (Amphibia: Leptodactylidae). Proceedings of the Biological Society of Washington, 125(3): 276-294. doi:10.2988/12-04.1

Heyer, Miriam M., Heyer, W. R. and de Sa, Rafael O. 2012. Leptodactylus pentadactylus. Catalogue of American Amphibians and Reptiles, 887: 1-48.

Heyer, W. R. and de Sá, Rafael O. 2011. Variation, Systematics, and Relationships of the Leptodactylus bolivianus Complex (Amphibia: Anura: Leptodactylidae). Smithsonian Contributions to Zoology, 635: 1-58. doi:10.5479/si.00810282.635.1

Laura Ponssa, Maria, Heyer, W. Ronald and De Sá, Rafael Omar 2010. Phylogeny of the genus Leptodactylus (Anura, Leptodactylidae) based on characters of morphology and mode of life. Cladistics, 26(2): 222

Heyer, W. R., Heyer, Miriam M. and De Sá, Rafael Omar 2010. Leptodactylus syphax. Catalogue of American Amphibians and Reptiles, 868: 1-9.

Heyer, W. R., Heyer, Miriam M. and De Sá, Rafael Omar 2010. Leptodactylus savagei. Catalogue of American Amphibians and Reptiles, 867: 1-19.

Furness, Andrew I., McDiarmid, Roy W., Heyer, W. Ronald and Zug, George R. 2010. Oviduct modifications in foam-nesting frogs, with emphasis on the genus Leptodactylus (Amphibia, Leptodactylidae). South American Journal of Herpetology, 5(1): 13-29.

Lavilla, Esteben O., Langone, Jose A., Caramaschi, Ulisses, Heyer, W. Ronald and De Sá, Rafael Omar 2010. The identification of Rana ocellata Linnaeus, 1758. Nomenclatural impact on the species currently known as Leptodactylus ocellatus (Leptodactylidae) and Osteopilus brunneus (Gosse, 1851)(Hylidae). Zootaxa, 2346: 1-16.

Camargo, Arley, Heyer, W. Ronald and De Sá, Rafael Omar 2009. Phylogeography of the frog Leptodactylus validus (Amphibia: Anura): Patterns and timing of colonization events in the Lesser Antilles. Molecular Phylogenetics and Evolution, 53: 571-579.

Heyer, W. R. and Barrio-Amorós, César Luis 2009. The advertisement calls of two sympatric frogs, Leptodactylus lithonaetes (Amphibia: Anura: Leptodactylidae) and Pristimantis vilarsi (Amphibia: Anura: Strabomantidae). Proceedings of the Biological Society of Washington, 122(3): 282-291.

Heyer, W. R. and Giaretta, Ariovaldo A. 2009. Advertisement calls, notes on natural history, and distribution of Leptodactylus chaquensis (Amphibia: Anura: Leptodactylidae) in Brasil. Proceedings of the Biological Society of Washington, 122(3): 292-305.

Heyer, Miriam H., Heyer, W. Ronald and De Sá, Rafael Omar 2009. Bibliography of the frogs of the Leptodactylus clade - Adenomera, Hydrolaetare, Leptodactylus, Lithodytes (Amphibia, Anura, Leptodactylidae). Volume 2. Indices. Smithsonian Herpetological Information Service, 2: 1-167.

Heyer, Miriam H., Heyer, W. Ronald and De Sá, Rafael Omar 2009. Bibliography of the frogs of the Leptodactylus clade - Adenomera, Hydrolaetare, Leptodactylus, Lithodytes (Ampbibia, Anura, Leptodactylidae). Volume 1. References. Smithsonian Herpetological Information Service, 1: 1-677.

Fernández, Miguel, Cole, Daniel, Heyer, W. Ronald, Reichle, Steffen and De Sá, Rafael Omar 2009. Predicting Leptodactylus (Amphibia, Anura, Leptodactylidae) distributions: Broad-ranging versus patchily distributed species using a presence-only environmental niche modeling technique. South American Journal of Herpetology, 4(2): 103-116.