| White-footed Voles inhabit a relatively small area of the Pacific coast of North America, from Humboldt Bay, California, north to the Columbia River. They have small eyes, and their claws are suitable for burrowing, so they probably have that habit, though it has not been observed. Most of the few specimens collected have been found near rivers or streams. Their fur is long, soft, and brownish; the tail is long and thinly haired, black above and white below; and the feet are white on top. These Voles breed year-round and have litters of 2-4 young.
Merriam, C.H., 1901. Proceedings of the Biological Society of Washington, 14:125.
Mammal Species of the World