Anatomical Evidence for Knuckle-walking Found
in Early Human Fossils

 

Summary of article appearing in Nature Vol. 404; 23 March 2000.

Article by Brian G. Richmond and David S. Strait


Several studies on the genetic information carried in our DNA and RNA point to chimpanzees being more closely related to humans than to gorillas. However, many researchers point out that chimpanzees and gorillas share several highly specialized anatomical traits with one another. These may indicate that a closer relationship exists between these two apes, with humans a slightly more distant evolutionary cousin.

These anatomical traits are mainly centered around adaptations for knuckle-walking. When gorillas and chimpanzees walk on the ground, they usually do so quadrupedally (on four legs). They support the weight of the front part of their body on the middle phalanges (the portion of the finger between the first and second knuckle of their hands, hence the term "knuckle-walking"). This is a highly specialized behavior that requires many modifications of the bones of the wrist, hand, and fingers to create a locking mechanism that is able to support their body weight. (Try doing a push-up on your middle phalanges!)

A serious question is then raised: If chimpanzees and humans are more closely related (as some genetic experiments suggest), did chimpanzees and gorillas evolve the modifications for knuckle-walking independently? To many paleontologists, this would seem a bit too coincidental.

Two researchers with the Smithsonian's Human Origins Program and George Washington University have recently identified evidence in early humans for a knuckle-walking ancestor. Using specimens of gorilla, chimpanzee, human and other primates that do not knuckle-walk, they were able to identify specific features of the radius, one of the bones in the forearm, that are indicative of knuckle-walking. Specifically they identified a ridge of bone that projects down into the wrist, stabilizing it when weight is placed on it in chimpanzees and gorillas.

Identifying a feature indicative of knuckle-walking at the end of the radius was important, because two early human species from near the beginning of human evolutionary history have preserved specimens of this bone: Australopithecus anamensis (KNM ER 20419) and Australopithecus afarensis (AL 288-1, the famous "Lucy" specimen). Upon examination of these specimens a clear ridge of bone was evident extending into the wrist from the radius. This ridge was not as pronounced as it is in modern chimpanzees and gorillas, which indicates that neither of these early human species were active knuckle-walkers. Rather, it suggests that these were traits inherited from a knuckle-walking ancestor. Examination of later-occurring early human species (Australopithecus africanus and Paranthropus robustus) showed that by three million years ago this trait had been lost in the human lineage.

This study implies that early humans evolved from an ancestor that was originally a knuckle-walker, and this raises the possibility that knuckle-walking evolved only once for gorillas, humans and chimpanzees. While we have stopped using this form of locomotion, the other African apes have retained it, and so they retain the similar anatomy that is associated with it.

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